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For example, excess copper in plants can form a hydroxyl radical from H 2 O 2 via the Haber-Weiss cycle ( Figure 3 3.4 Impact of exogenous ergothioneine on the pyridine dinucleotide profiles of stored broccoli NADPH is required as a cofactor for many metabolic processes in plants, including the major ROS-generating enzyme NADPH-dependent oxidase activity, as well as the electron donor-driving GR activity in the ascorbate-GSH recycling pathway ( Figure 4 )