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In their case they calculated the average number of mutations per culture 'from the proportion of cultures containing no resistant bacteria at the moment of the test' in accordance with Poisson's law (Luria, Delbrck)

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doi: 10.1073/pnas.1318431110 147

Eur J Pharmacol 583(1):7383 di Maio R, Mastroberardino PG, Hu X et al (2013) Thiol oxidation and altered NR2B/NMDA receptor functions in in vitro and in vivo pilocarpine models: implications for epileptogenesis [J]