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The copper ion is integral to GHK-Cu's biological activity, serving as a cofactor in enzymatic reactions involved in collagen crosslinking and antioxidant defence mechanisms

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The specific kinases involved in occludin phosphorylation remain elusive

The mainstream copper-biology research community and the broader peptide research community have engaged with the GHK-Cu literature at various points, but the depth of independent characterization outside the Pickart-affiliated research has been variable