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This response could be caused by an increase in ROS or ammonia, which aged explants cannot process as effectively due to lower GSH production

10.1039/c5tb02174j 73 HanF.ChenQ.SuJ.ZhengA.ChenK.SunS.et al (2019)

These results suggest that metformin promotes GLUT1 expression through TXNIP suppression, independently of AMPK, and probably via mitochondrial complex I inhibition