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All structure representations in the manuscript were generated using UCSF Chimera 71 , ChimeraX 74 , and PyMOL 75

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332 HOTAIR suppresses tumor suppressor genes like PGR, PCDH10, PCDHB5, and JAM2, promoting breast cancer development, and is overexpressed in colorectal, hepatocellular, gastrointestinal, and NSCLC

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