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Precipitated histones were pelleted at 10,000 x g for 5 minutes, washed once with 0.1% HCl in acetone and twice with acetone followed by centrifugation at 14,000 x g for 5 minutes

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While cationic polymers like PEI can enhance loading, they introduce well-documented cytotoxicity constraints, forcing an explicit engineering trade-off between delivery potency and biological safety (Shirbaghaee and Bolhassani, 2016)