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Abbreviations GABA: Gamma-aminobutyric acid ATP: Adenosine triphosphate ADP: Adenosine diphosphate CARNS1: Carnosine synthase 1 ATPGD1: ATP-grasp domain-containing protein CN1: Serum carnosinase CN2: Tissue carnosinase CNDP1: Carnosine dipeptidase 1 CNDP2: Carnosine dipeptidase 2 DHPR: Dihydropyridine receptor SERCA: Sarcoendoplasmic reticulum calcium ATPase ROS: Reactive oxygen species HNE: 4-Hydroxy-2-nonenal MDA: Malondialdehyde ALE: Advanced lipoxidation end products NADPH: Nicotinamide adenine dinucleotide phosphate NF-B: Nuclear factor kappa B TNBS: 2,4,6-Trinitrobenzene sulfonate TNF-: Tumor necrosis factor alpha NOS: Nitric oxide synthase NO: Nitric oxide IFN-: Interferongamma TGF-: Transforming growth factor- CCL24: Chemokine (CC motif) ligand 24 Smad3: Mothers against decapentaplegic homolog 3 ALK5: Activin receptor-like kinase-5 Smad2: Mothers against decapentaplegic homolog 2 SV40: Simian virus 40 C-6: Rat glioma cell line IL-6: Interleukin-6 IL-8: Interleukin 8 H 2 O 2 : Hydrogen peroxide LCMS/MS: Liquid chromatography-tandem mass spectrometry ISRCTN: International Standard Randomised Controlled Trial Number References Abe H (2000) Role of histidine-related compounds as intracellular proton buffering constituents in vertebrate muscle

CSCs reportedly exhibit higher antioxidant gene expression levels and lower ROS levels than normal cells 151

Vaping exerts a detrimental effect on the airway epithelium, leading to significant cellular damage in the lungs and disruption of mucociliary clearance, both of which are important in the normal functioning and health of the lungs and airways

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