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In short, organoids were incubated overnight at 4 C with primary antibodies binding ZO-I (Thermo Fisher Scientific catalog number 61-7300, RRID: AB_2533938), ITGB1 gene product (R and D Systems catalog number MAB1778, RRID: AB_357909) and Ki67 (Millipore catalog number AB9260, RRID: AB_2142366) or ITGA2 -encoded gene (Abcam catalog number ab30487, RRID: AB_775702)